The Program in Neuroscience draws together neuroscientists from across Harvard. The physical home base of the program is located at the Longwood Campus of Harvard Medical School, in the Department of Neurobiology.

Research sites include the Longwood Medical Area, Cambridge Campus, Massachusetts General Hospital, and the McLean Hospital. The Center for Brain Science unites many neuroscience labs and houses in the newly established Swartz Program in Theoretical Neuroscience.